Cassidinae Checklist: Tortoise Beetles

13 August 2019

Tortoise beetles belong to the subfamily Cassidinae. They are characterised by a broad domed oval shaped elytra, like that of a tortoise shell. The larvae are flattened and spiny, often with a swollen “tail” which is an accumulation of their excrement and shedded exoskeletons.

Most species in this page were identified with the kind assistance of Michael Geiser, Dominik Hofer and Rob Westerduijn on Facebook.

All photos are of live subjects shot in the field, with the dorsal view selected where available. Click on individual photos for larger views and views from other angles. As specimens were not collected, identifications were done purely based on photographs and may not be 100% accurate.

Some useful references:

This page will be updated regularly, please let me know if you spot any mistakes.

View my complete Flickr photo set: Cassidinae – Tortoise Beetles.

Region Filters

Click on any region to show only records from that region.

Class: Insecta Linnaeus, 1758

Order: Coleoptera Linnaeus, 1758

Family: Chrysomelidae Latreille, 1802

Subfamily: Cassidinae Gyllenhal, 1813

Tribe: Aspidimorphini Chapuis, 1875

Tortoise beetle (Aspidimorpha deusta) - P9168221

Singapore: Aspidimorpha deusta (Fabricius, 1775)

Tortoise Beetle (Aspidimorpha miliaris) - DSC_1875

Singapore: Aspidimorpha miliaris (Fabricius, 1775)

Golden tortoise beetle (Aspidimorpha furcata) - DSC_7167

East Malaysia: Aspidimorpha cf. furcata (Thunberg, 1789)

Golden tortoise beetle (Aspidimorpha cf. furcata) - P5076799

Singapore: Aspidimorpha cf. furcata (Thunberg, 1789)

Tortoise beetle (Laccoptera nepalensis) - DSC_5063

Singapore: Laccoptera nepalensis (Boheman, 1855)

Golden tortoise beetle (Aspidimorpha cf. ertli) - DSC_2776

Mozambique: Aspidimorpha cf. ertli Spaeth, 1906

Golden tortoise beetle (Aspidimorpha sp.) - PB192852

Thailand: Aspidimorpha sp.

Tribe: Basiprionotini Gressitt, 1952

Tortoise beetle (Basiprionota sp.) - PB214807

Thailand: Basiprionota sp.

Tribe: Cassidini Gyllenhal, 1813

Tortoise beetle (Cassida sp.) - PB182160

Thailand: Cassida sp.

Tortoise beetle (Rhacocassis flavoplagiata) - P3127178

East Malaysia: Rhacocassis flavoplagiata (Baly, 1863)

Tortoise beetle (Cassida sp.) - P5230122

Singapore: Cassida cf. circumdata (Herbst, 1799)

Tortoise Beetle (Cassida sp.) - DSC_4434

Thailand: Cassida sp.

Tortoise beetle (Chiridopsis cf. scalaris) - P5076744

Singapore: Chiridopsis cf. scalaris (Weber,1801)

Tortoise beetle (Ischnocodia annulus) - DSC_9301

Belize: Ischnocodia annulus (Fabricius, 1781)

Tortoise beetle (Ischnocodia annulus) - DSC_8049

Belize: Ischnocodia annulus (Fabricius, 1781)

Curry leaf tortoise beetle (Silana farinosa) - DSC_7449

Singapore: Silana farinosa (Boheman, 1856)

Tortoise beetle (Thlaspida biramosa) - DSC_9348

West Malaysia: Thlaspida biramosa (Boheman, 1855)

Tortoise beetle (Malayocassis hilaris) - P7084460

East Malaysia: Malayocassis hilaris (Boheman, 1855)

Tortoise beetle (Charidotis ambigua) - P6078720

Ecuador: Charidotis ambigua Boheman, 1855

Target tortoise beetle (Plagiometriona phoebe) - P6100575

Ecuador: Plagiometriona phoebe (Boheman, 1855)

Tortoise beetle (Cassidini) - P6067305

Ecuador: Cassidini P6067305

Golden tortoise beetle (Hybosa sp.) - P6088928

Ecuador: Hybosa sp.

Tribe: Hemisphaerotini Monrós & Viana, 1951

Mottled tortoise beetle (Spaethiella sp.) - DSC_2789

Peru: Spaethiella sp.

Tribe: Imatidiini Chapuis, 1875

Tortoise beetle (Aslamidium bolivianum) - DSC_3221

Peru: Aslamidium cf. bolivianum Borowiec & Sassi, 2001

Tribe: Notosacanthini Hincks, 1952

Tortoise Beetle (Notosacantha cf. rufa) - DSC_4573

West Malaysia: Notosacantha cf. rufa (Wagener, 1881)

Tribe: Oncocephalini Chapuis, 1875

Leaf-mining beetle (Oncocephala sp.) - DSC_7681

Singapore: Oncocephala sp.

Tribe: Physonotini Spaeth, 1942

Tortoise beetle (Platycycla deruta) - DSC_9135

Belize: Platycycla deruta Boheman, 1854

Tribe: Sceloenoplini Uhmann, 1930

Leaf beetle (Sceloenopla sp.) - P6099797

Ecuador: Sceloenopla sp. P6099797

Leaf beetle (Sceloenopla sp.) - P6111603

Ecuador: Sceloenopla sp. P6111603

Tribe: Spilophorini Chapuis, 1875

Tortoise beetle (Calyptocephala cf. gerstaeckeri) - DSC_0452

Belize: Calyptocephala cf. gerstaeckeri Boheman, 1862

Unidentified Cassidinae

Leaf beetle (Cassidinae) - P6154363

Ecuador: Cassidinae P6154363

Tortoise beetle (Cassidinae) - PB215399

Thailand: Cassidinae PB215399


Tortoise beetle eggs? (Silana farinosa) - DSC_9502

Singapore: Silana farinosa (Boheman, 1856)


Tortoise Beetle larva (Silana farinosa) - DSC_5086

Singapore: Silana farinosa (Boheman, 1856)

Curry leaf tortoise beetle larvae (Silana farinosa) - DSC_9431

Singapore: Silana farinosa (Boheman, 1856)

Tortoise beetle larva (Cassidinae) - ESC_0002

Singapore: Unidentified

Tortoise beetle larva (Aspidimorpha cf. miliaris) - DSC_4584

Singapore: Aspidimorpha cf. miliaris (Fabricius, 1775)

Tortoise beetle larva (Aspidimorpha deusta) - P9168210

Singapore: Aspidimorpha deusta (Fabricius, 1775)


Tortoise Beetle pupa (Silana farinosa) - DSC_8136

Singapore: Silana farinosa (Boheman, 1856)

Tortoise beetle larva (Aspidomorpha sp.) - P5230058

Singapore: Aspidimorpha sp.

Tortoise beetle pupa (Aspidimorpha deusta) - P9168241

Singapore: Aspidimorpha deusta (Fabricius, 1775)

  1. Reply

    Pedro Castillo carrillo

    13 April 2020


  2. Reply

    Nate Sorensen

    6 May 2022

    Hello Nicky you have a nice arrangement of Beatles here! I found one today that I cannot find anywhere on Google I was wondering if you had an idea it’s a type of Is tortoise beetle but I have not seen anything exactly like it on Google. If you would like to email me back to see a picture dear picture I could send it to you.. Thank you…nate



Hi my name is Nicky Bay. I am a macro photographer, instructor and book author, travelling the world to document the vast micro biodiversity that nature has to offer. Follow my updates and discover with me the incredible beauty and science behind our planet's micro creatures!

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